In 1951, mathematician Alan Turing is questioned by police after an apparent home break-in, and he obliquely refers to his work at Bletchley Park during World War II. In 1928, Turing, constantly bullied at boarding school, befriends Christopher Morcom, who sparks his interest in cryptography. Turing develops romantic feelings, but Christopher dies of bovine tuberculosis before he can confess. When Britain declares war on Germany in 1939, Turing is an established mathematician. He joins the cryptography team of Hugh Alexander, John Cairncross, Peter Hilton, Keith Furman, and Charles Richards in Bletchley Park, directed by Commander Alastair Denniston. Their main employment is analysis of the Enigma machine the Wehrmacht uses to send coded messages. Difficult to work with, and believing his colleagues to be inferior, Turing works alone to design a deciphering machine, which he names "Christopher". When Denniston refuses to fund the £100,000 construction cost, Turing contacts Prime Minister Winston Churchill who appoints him as team leader and provides the funds. Turing then fires Furman and Richards and places a difficult crossword in newspapers as a test to find replacements. Cambridge graduate Joan Clarke passes Turing's test but her family refuses her permission to work with the male cryptographers. Turing arranges for her to live and work with the women who intercept the messages, and shares his plans with her. Clarke helps Turing warm to the others, who begin to respect him. Turing's machine is constructed but cannot determine the Enigma encryption settings quickly enough, as the Germans reset them each day. Denniston orders it destroyed and Turing fired, and the other cryptographers threaten to leave. When Clarke plans to quit because of her parents, Turing proposes, which she accepts. During their engagement party, Turing confirms his homosexuality to Cairncross, who advises him to keep it a secret. Overhearing a clerk talking about messages received from the same German coder, Turing has an epiphany: he can program the machine to decode words he knows exist in certain messages. The theory is proven correct when the German coder consistently opens messages with a standard plaintext German script, which reveals enough of the daily Enigma code to permit decoding of that day's messages. The cryptographers celebrate this breakthrough. Discovering through decoded intercepts that a convoy is in danger of a German attack, Turing fears a sudden response will notify the Germans that Enigma is compromised and change it. The team concludes they cannot act on every decoded message, and they issue no warning about the attack on the convoy, even though Peter, whose brother is serving in the convoy, begs them. Turing creates a statistical model to select which intelligence to act on, to maximize effect on the enemy while minimizing the risk of German suspicion. Turing confronts Cairncross, who he has uncovered as a Soviet spy. Cairncross argues the Soviets are allies working for the same goals, and threatens to retaliate by disclosing Turing's sexuality. When MI6 agent Stewart Menzies appears to threaten Clarke, Turing shares his revelation about Cairncross. Menzies reveals in turn that he already knew, and is using Cairncross to leak misinformation to the Soviets for British benefit. Turing urges Clarke to leave Bletchley Park and admits his sexuality. She admits always being suspicious but insists they would have been happy together anyway. Fearing for her safety, Turing replies he never cared for her, and only used her for her cryptography skills. The heartbroken Clarke stays on, defying her parents and Turing. After the war, Menzies has the cryptographers destroy all traces of the project, as MI6 wants foreign governments to feel secure using their own code machines without fear of British interception. The team is instructed never to meet again or share what they have done. In 1952, Turing is convicted of gross indecency and undergoes chemical castration to be spared from prison so he can continue his work. Clarke visits him, witnesses his physical and mental deterioration, and tries to comfort him. The epilogue shows Turing committed suicide on June 7, 1954, after a year of government-mandated hormonal therapy. In 2013, Queen Elizabeth II granted him a posthumous Royal Pardon. Historians estimate that breaking Enigma shortened the war by over 2 years, saving over 14 million lives. Turing's work is now recognized as an essential step toward the development of modern computers.

Dr. John Mathewson discovers a new process for refining plutonium to purities greater than 99.997 percent. The United States government provides him a laboratory located in Ithaca, New York, masked as a medical company. John moves to Ithaca and meets real estate agent Elizabeth Stephens while searching for an apartment. He attempts to win the affections of the single mother by inviting her teenage son Paul to take a tour of the lab. John is confident in the lab's cover story but Paul, an unusually gifted student with a passion for science, becomes suspicious when he discovers a statistically impossible patch of five-leaf clover on the grounds. Paul and his aspiring journalist girlfriend, Jenny Anderman, decide to expose the weapons factory to the media, stealing a container of plutonium as evidence. Once they succeed, Paul declares that the plutonium alone is insufficient, as no-one would be impressed by two kids stealing something from a lab. Instead, he convinces Jenny he can create the world's first privately built nuclear device, exposing the lab to the world by audaciously entering it into the New York Science Fair. After convincing his mother and his school that his project is about hamsters bred in darkness, he begins research and construction of the bomb. The lab discovers that a container of plutonium has been replaced by a bottle of shampoo mixed with glitter. A military investigation team, led by Lt. Colonel Conroy, arrives on the scene and determines that Paul is responsible for stealing the plutonium. Suspecting him of terrorist intent, the investigators search Paul's home and discover that he and Jenny have left for the science fair. After the agents capture the couple in New York City, John, who feels personally responsible for the crisis, talks privately with Paul and persuades him to give the bomb to the agents before a group of other participants at the science fair help Paul and Jenny escape from the hotel. In an effort to expose the lab, Paul hatches a plan to return the bomb on his own terms. Ensuring Jenny is a safe distance away, he calls the agents from a pay phone and walks into the lab with the bomb while being surrounded by snipers and agents. During the standoff, negotiations stall and Paul arms the bomb. John, convinced that Paul is not an actual terrorist, attempts to intercede on his behalf. Due to radiation from the plutonium, the bomb's timer activates on its own and begins to count down with increasing speed. Paul suggests taking the bomb to a quarry outside of town, but John admits that he had not fully understood the ramifications of his plutonium refining process. As a result, Paul's bomb will have a nuclear blast yield nearly five times larger than the one that destroyed Hiroshima if it detonates. Desperate to defuse the bomb, all sides put down their weapons and frantically work as a team to dismantle it. They manage to disarm the bomb a fraction of a second before it explodes. After a brief moment of relief, Conroy decides to arrest Paul. John refuses to cooperate and opens the door to the lab, revealing a large crowd, including Jenny and the press. The film ends with Paul freely departing the scene.

In 2019, Lincoln Six Echo and Jordan Two Delta live with others in an isolated compound. This dystopian community is governed by a strict set of rules. The residents are told that the outside world has become too contaminated to support life with the exception of a pathogen-free island. Each week, one resident gets to leave the compound and live on the island by way of a lottery. Lincoln begins having dreams that he knows are not from his own experiences. Dr. Merrick, a scientist who runs the compound, is concerned so he places probes in Lincoln's body to monitor his cerebral activity. While secretly visiting the off-limits power facility in the basement where technician James McCord works, Lincoln discovers a live moth in a ventilation shaft, leading him to deduce the outside world is not really contaminated. Lincoln follows the moth to another section, where he discovers the "lottery" is actually a system to selectively remove inhabitants from the compound, where the "winner" is then used for organ harvesting, surrogate pregnancies, and other important purposes for each one's wealthy sponsor, of whom they are clones. Merrick learns Lincoln has discovered the truth about his existence, which compels Lincoln to escape. Meanwhile, Jordan has been selected for the island. She and Lincoln escape the facility and emerge in the desert. He explains the truth to Jordan, and they set out to discover the real world. Merrick hires Burkinabé mercenary and former GIGN operative Albert Laurent to find and return them to the compound. Lincoln and Jordan find McCord in a bar, who explains that all the facility residents are clones of wealthy sponsors kept ignorant about the real world and conditioned to never question their environment or history. Merrick explains to Laurent that while the public is told that the clones are kept in a persistent vegetative state, trials had shown that the organs could only survive if the clones had consciousness. McCord provides the name of Lincoln's sponsor, yacht designer Tom Lincoln, in Los Angeles and helps them to the Yucca maglev station, where they board an Amtrak train to Los Angeles before mercenaries kill him. In New York City, Jordan's sponsor, supermodel Sarah Jordan, is comatose following a car crash and requires transplants from Jordan to survive. Lincoln and Jordan evade both the Los Angeles Police Department and the mercenaries and arrive at Tom's house. Lincoln meets Tom, who gives him some explanation about the cloning institute, causing Lincoln to realize that he has gained Tom's memories. Tom seemingly agrees to help Lincoln and Jordan reveal Merrick's crimes to the public, but secretly betrays them to Merrick and Laurent, as he desperately needs Lincoln's liver to survive his cirrhosis. Tricking Lincoln into leaving with him, although Jordan had warned him she believed he was lying, Tom brings him to an ambush that results in a car chase through suburban Los Angeles. It ends with Lincoln tricking Laurent into believing Tom is the clone and killing him, allowing him to assume Tom's identity. Returning to Tom's home, Lincoln and Jordan give in to their romantic urges and have sex. Merrick surmises that a cloning defect was responsible for Lincoln's memories and behavior, resulting in him and every future clone generation to question their environment and even tap into their sponsor's memories. To prevent this, he decides to eliminate the four latest generations of clones. Lincoln and Jordan, however, plan to liberate the other clones. Posing as Tom, Lincoln returns to the compound to destroy the holographic projectors that conceal the outside world. Jordan allows herself to be caught to assist Lincoln's plan. Laurent, who has moral qualms about the clones' treatment after witnessing their fight for survival and learning that Sarah may not survive even with the organ transplants, helps Jordan. Lincoln kills Merrick with a harpoon gun, and the clones are freed, seeing the outside world for the first time. As Laurent seemingly gives up his mercenary life, Lincoln and Jordan sail away in one of Tom's boats together toward an island, fulfilling their dream of one day going to such a place.

In 1947, over the Muroc Army Air Field in California, a number of test pilots are killed while flying high-speed aircraft such as the rocket-powered Bell X-1. After another pilot, Slick Goodlin, demands $150,000 (equivalent to $2,163,000 in 2025) to attempt to break the sound barrier, war hero Captain Chuck Yeager receives the chance to fly the X-1. Yeager becomes the first person to fly at supersonic speed, defeating the "demon in the sky." Six years later, Muroc, now Edwards Air Force Base, still attracts the best test pilots. Yeager (now a major) and friendly rival Scott Crossfield repeatedly break each other's speed records. They often visit the Happy Bottom Riding Club run by Pancho Barnes, who classifies the pilots at Edwards as either "prime" (such as Yeager and Crossfield) that fly the best equipment or newer "pudknockers" who only dream about it. Gordon "Gordo" Cooper, Virgil "Gus" Grissom and Donald "Deke" Slayton, captains of the United States Air Force, are among the "pudknockers" who hope to also prove that they have "the Right Stuff." The tests are no longer secret, as the military soon recognizes that it needs good publicity for funding. Cooper's wife, Trudy, and other wives are afraid of becoming widows but cannot change their husbands' ambitions and desire for success and fame. In 1957, the launch of the Soviet Sputnik satellite alarms the United States government. Politicians such as Senator Lyndon B. Johnson and military leaders demand that NASA help America defeat the Soviets in the new Space Race. The search for the first Americans in space excludes Yeager because he lacks a college degree. Grueling physical and mental tests select the Mercury Seven astronauts, including John Glenn of the United States Marine Corps, Alan Shepard, Walter Schirra and Scott Carpenter of the United States Navy, as well as Cooper, Grissom and Slayton; they immediately become national heroes. Although many early NASA rockets explode during launch, the ambitious astronauts all hope to be the first in space as part of Project Mercury. Although engineers see the men as passengers, the pilots insist for the Mercury spacecraft to have a window, a hatch with explosive bolts, and pitch-yaw-roll controls. However, the Soviet Union beats them into space on April 12, 1961, with the launch of Vostok 1 carrying Yuri Gagarin. The seven astronauts are determined to match and to surpass the Soviets. Shepard is the first American to reach space on the 15-minute sub-orbital flight of Mercury-Redstone 3 on May 5. After Grissom's similar flight of Mercury-Redstone 4 on July 21, the capsule's hatch blows open and quickly fills with water. Grissom escapes, but the spacecraft, overweight with seawater, sinks. Many criticize Grissom for possibly panicking and opening the hatch prematurely. Glenn becomes the first American to orbit the Earth on Mercury-Atlas 6 on February 20, 1962; he survives a possibly loose heat shield and receives a ticker-tape parade. He, his colleagues, and their families become celebrities, including a gigantic celebration in the Sam Houston Coliseum to announce the opening of the Manned Spacecraft Center in Houston, despite the fear of Glenn's wife, Annie, of public speaking because of a stutter. Although test pilots at Edwards mock the Mercury program for sending "spam in a can" into space, they recognize that they are no longer the fastest men on Earth, and Yeager states that "it takes a special kind of man to volunteer for a suicide mission, especially when it's on national TV." While testing the new Lockheed NF-104A, Yeager attempts to set a new altitude record at the edge of space but is nearly killed in a high-speed ejection when his engine fails. Though seriously burned after reaching the ground, Yeager gathers up his parachute and walks to the ambulance, which proves his worth. On May 15, 1963, Cooper has a successful launch on Mercury-Atlas 9, which ends the Mercury program. As the last American to fly into space alone, he "went higher, farther, and faster than any other American... for a brief moment, Gordo Cooper became the greatest pilot anyone had ever seen."

In 1981, San Francisco salesman Chris Gardner invests his entire life savings in portable bone-density scanners, which he demonstrates to doctors and pitches as a handy improvement over standard X-rays. The scanners play a vital role in Chris's life. While he can sell most of them, the time lag between the sales and his growing financial demands enrages his wife, Linda, who works as a hotel maid. The economic instability increasingly erodes their marriage, despite caring for Christopher Jr., their soon-to-be five-year-old son. While out on a trip to sell one of his last scanners, Chris meets Jay Twistle, a lead manager and partner for Dean Witter Reynolds and impresses him by solving a Rubik's Cube during a taxi ride. After Jay leaves, Chris skips out on paying the fare as he lacks the money, causing the driver to angrily chase him into a BART station. Despite boarding a train, Chris loses one of his scanners in the process. However, Chris' new relationship with Jay earns him the chance to become an intern stockbroker. The day before the interview, Chris begrudgingly agrees to paint his apartment for free to postpone eviction by his landlord for late rent. However, two policemen arrive to arrest Chris for multiple unpaid parking tickets. Chris has to spend the night in jail, complicating his schedule for the interview the next day. Chris narrowly arrives at Dean Witter's office on time, albeit still in shabby, paint-splattered clothes. Despite his appearance, Chris still impresses the interviewers and lands a six-month unpaid internship. He is among 20 interns competing for a paid position as a stockbroker. Chris' unpaid internship does not please Linda, who leaves for New York since she might get a job at her sister's boyfriend's new restaurant. After Chris tells Linda she is incapable of being a single parent, she leaves Christopher in Chris' care. However, Chris is further set back when his already diminished bank account is garnished by the IRS for unpaid income taxes, and Chris and Christopher are evicted. With only $21.33 in his bank account, Chris and Christopher are left homeless and desperate; Chris is able to get a motel room, but the locks are then changed when he is unable to pay on time. They are forced at one point to stay in a restroom at a BART station. Other days, Chris and Christopher spend nights at a homeless shelter, in BART, or, if Chris manages to procure sufficient cash, at a hotel. Later, he finds the bone scanner that he lost in the BART station earlier. However, Chris found it damaged, yet he still manages to repair it by selling his own blood for money availability and after that sells it to a physician, thus completing all sales of his scanners. Disadvantaged by his limited work hours and knowing that maximizing his client contacts and profits is the only way to earn the broker position, Chris develops several ways to make sales calls more efficiently, including reaching out to potential high-value customers in person, a violation of firm protocol. One sympathetic prospect, Walter Ribbon, a top-level pension fund manager, even takes Chris and Christopher to a San Francisco 49ers game, where Chris meets some of Mr. Ribbon's friends, who are also potential clients. Regardless of his challenges, Chris never reveals his lowly circumstances to his colleagues, even going so far as to lend one of his supervisors, Mr. Frohm, five dollars in his wallet for cab fare. Chris also studies for and aces the stockbroker license exam although he doubts that he did well. As Chris concludes his last day of internship, he is summoned to a meeting with the partners. Mr. Frohm lets Chris know that he has won the coveted full-time position and reimburses Chris for the previous cab ride. Fighting back tears, he shakes hands with the partners, then rushes to Christopher's daycare to embrace him. They walk down a street and joke with each other. An epilogue reveals that Chris went on to form his own multimillion-dollar brokerage firm in 1987 and sold a minority stake in it in a multi-million-dollar deal in 2006.

In intercut frame stories, two ongoing depositions play out: in one, Mark Zuckerberg is being sued by his former friend Eduardo Saverin, and in the other, he is sued by twins Cameron and Tyler Winklevoss. On October 28, 2003, 19-year-old Zuckerberg, a Harvard University sophomore, is dumped by his girlfriend, Erica Albright. Returning to his dorm, Zuckerberg writes an insulting post about her on his LiveJournal blog. Zuckerberg creates a campus website called Facemash by hacking and downloading photos of female students from house face books, then allowing site visitors to rate their attractiveness. After traffic to the site crashes parts of Harvard's computer network, Zuckerberg is given six months of academic probation. Facemash's popularity attracts the attention of the Winklevoss twins and their business partner Divya Narendra. The trio invites Zuckerberg to work on Harvard Connection, a social network exclusive to Harvard students focused on dating. Soon after, Zuckerberg approaches Saverin with an idea for " TheFacebook ", a social networking website that would be exclusive to Ivy League students. As its CFO, Saverin provides $1,000 in seed funding to allow Zuckerberg to build the website, which quickly becomes popular. When they learn of TheFacebook, the Winklevoss twins and Narendra are incensed, believing that Zuckerberg stole their idea while misleading them by stalling development on the Harvard Connection. They raise their complaint with Harvard President Larry Summers, who is dismissive and sees no value in disciplinary action on TheFacebook or Zuckerberg. Saverin and Zuckerberg meet fellow student Christy Lee, who asks them to "Facebook me," a phrase that impresses them. As TheFacebook grows in popularity, Zuckerberg expands the network to Yale, Columbia, and Stanford. Saverin begins dating Lee, who arranges for him and Zuckerberg to meet Napster co-founder Sean Parker. Parker presents a "billion-dollar" vision for the company, impressing Zuckerberg. Saverin dismisses Parker as paranoid and delusional, except for his suggestion to rename TheFacebook to Facebook. Saverin deposits $18,000 into a new account as Zuckerberg relocates the company to Palo Alto on Parker's advice; Saverin remains in New York to work on business development while breaking up with Lee. Parker later moves into the house that Zuckerberg is using as a base of operations and becomes more involved with the company, much to Saverin's annoyance. After narrowly losing in the 2004 Henley Royal Regatta for Harvard against the Hollandia Roeiclub, the Winklevoss twins discover that, through Parker, Facebook has expanded to Europe with Oxford, Cambridge and LSE, and decide to sue the company for intellectual property theft. Meanwhile, Saverin objects to Parker's decision-making for Facebook and freezes the company's bank account in the resulting dispute. He relents when Zuckerberg reveals that they have secured $500,000 from angel investor Peter Thiel. Saverin is invited to the company's new headquarters in San Francisco on the pretense of attending a business meeting and "millionth user party," but is enraged when he discovers that the new investment deal dilutes his share of Facebook from 34% to 0.03%, without diluting the ownership percentage of any other owner. Furthermore, he is removed from the masthead as co-founder and CFO. He confronts Zuckerberg and Parker, vowing to sue as security removes him from the building. Shortly after, Parker is apprehended for cocaine possession with a minor at a party celebrating one million users. He attempts to blame Saverin, prompting Zuckerberg to cut ties with him. Marylin Delpy, a junior lawyer for Zuckerberg, informs Zuckerberg that they will settle with Saverin since the sordid details of Facebook's founding and Zuckerberg's callous attitude will make him unsympathetic to a jury. Alone, Zuckerberg sends a Facebook friend request to Albright and repeatedly refreshes the page. Closing text reveals both cases were settled out of court and Zuckerberg became the world's youngest billionaire.

Filmmaker Martin "Marty" Di Bergi is filming a documentary about English rock band Spinal Tap 's 1982 United States concert tour to promote their new album, Smell the Glove. The band comprises childhood friends David St. Hubbins and Nigel Tufnel on vocals and guitar, along with bassist Derek Smalls, keyboardist Viv Savage, and drummer Mick Shrimpton. The documentary shows Spinal Tap's early days as the skiffle group The Originals; they renamed themselves the New Originals when it was discovered another band was already called The Originals, only to change it back when the original Originals changed their name to The Regulars. They later had a hit as the Thamesmen, "Gimme Some Money", before changing their name to Spinal Tap and achieving a hit with the flower power anthem "Listen to the Flower People"; they subsequently began performing heavy metal. Several of the band's previous drummers died under strange circumstances: John "Stumpy" Pepys died in a "bizarre gardening accident" that police said was better left unsolved, Eric "Stumpy Joe" Childs died choking on someone else's vomit, and Peter "James" Bond exploded on stage. Nigel shows Marty his extensive guitar collection (including a Fender Bass VI so valuable it cannot even be looked at), as well as a custom-made amplifier with volume knobs that go up to eleven; Nigel claims that this makes the amplifier "one louder" than most others, on which the volume setting only goes up to "ten". Tensions rise between the band and their manager, Ian Faith, as several shows are canceled due to low ticket sales and major retailers refuse to sell Smell the Glove because of its sexist cover art. David's girlfriend Jeanine, a yoga and astrology devotee, joins the group on tour and participates in band meetings. Nigel and Ian dislike Jeanine's ideas for Spinal Tap's costumes and stage presentation. Without consulting the band, the band's record label releases Smell the Glove with an entirely black album cover. Despite Ian's assertion that it could have a similar appeal to the Beatles ' White Album, Smell the Glove fails to sell. Nigel suggests staging a lavish, Druid -themed stage show and asks Ian to order a replica Stonehenge trilithon. However, Nigel mislabels its dimensions, and the resulting prop is only 18 inches (46 cm) high rather than 18 feet (5.5 m), making the group a laughing stock. The group blames Ian, and when David suggests Jeanine should co-manage the group, Ian quits. The tour continues, rescheduled for much smaller venues, and Jeanine and David increasingly marginalize Nigel. At a gig at a United States Air Force base, Nigel is upset by an equipment malfunction and quits mid-performance. At their next gig, in an amphitheater at an amusement park where the band is billed below a puppet show, the band finds their repertoire is severely limited without Nigel. At Derek's suggestion, the band improvises an experimental "Jazz Odyssey", which is poorly received. On the last day of the tour, David and Derek consider ending Spinal Tap and exploring other projects, such as a musical about Jack the Ripper called Saucy Jack. Before they go on stage, Nigel arrives and tells them that Spinal Tap's song "Sex Farm" has become a major hit in Japan and that Ian wants to arrange a tour there. David bitterly refuses; later, however, as Nigel watches the band's performance from backstage, David relents and invites him onstage, delighting the crowd but infuriating Jeanine. Mick subsequently explodes on stage. Ian is rehired as the group's manager, and Spinal Tap (now with Joe "Mama" Besser as their drummer) performs a series of sold-out shows in Japan.

Leading software engineer Kevin Flynn, formerly employed by technology corporation ENCOM, now runs a video game arcade, where he hacks into ENCOM's system with a program called CLU, hoping to find proof that he is the true author of ENCOM's best-selling videogame, 'Space Paranoids'. However, ENCOM's Master Control Program (MCP) halts his progress and deletes CLU. Within ENCOM, programmer Alan Bradley and his girlfriend, engineer Lora Baines, discover that the MCP has closed off their access to projects. When Alan confronts the senior executive vice president, Ed Dillinger, he asserts the security measures are an effort to stop the hacking attempts. However, when Dillinger privately questions the MCP through his computerized desk, he realizes the MCP has expanded into a powerful virtual intelligence and has been illegally appropriating personal, business, and government programs to increase its own capabilities. As Dillinger rose to the top of ENCOM by presenting Flynn's games as his own, the MCP blackmails Dillinger by threatening to expose his plagiarism should he not comply with its directives. Lora deduces that Flynn is the hacker, and she and Alan go to his arcade to warn him. Flynn reveals that he has been trying to locate evidence proving Dillinger's guilt. Together, the three form a plan to break into ENCOM and unlock Alan's "Tron" program, a self-governing security measure designed to protect the system and counter the functions of the MCP. Once inside ENCOM, the three split up, and Flynn comes into direct conflict with the MCP through a laboratory terminal. Before Flynn can get the information he needs, the MCP uses an experimental laser to digitize and upload him into the ENCOM gaming grid. There, computer programs are living entities appearing in the likeness of the human "Users" (programmers) who created them. The space is ruled by the MCP and its second-in-command, Sark (an avatar of Dillinger), who coerce programs to renounce their belief in the Users and force those who resist to compete in deadly games. Flynn is put into the games and plays well; between matches, he befriends two other captured programs, Ram and Tron. The three escape into the system during a round of Light Cycle (an arcade game Flynn created and is skilled at), but Flynn and Ram become separated from Tron by an MCP pursuit party. While attempting to help a badly injured Ram, Flynn learns that he can manipulate portions of the system by accessing his programmer knowledge. Just before Ram "derezzes" (essentially, being erased), he recognizes Flynn as a User, and encourages him to find Tron and free the system. Using his newfound ability, Flynn rebuilds a broken recognizer and disguises himself as one of Sark's soldiers. Tron enlists help from Yori, a sympathetic program and avatar of Lora, and at an I/O tower receives information from Alan necessary to destroy the MCP. Flynn rejoins them, and the three board a hijacked solar sailer to reach the MCP's core. However, Sark's command ship derezzes the sailer, capturing Flynn and Yori and presumably killing Tron. Sark leaves the command ship and orders its deresolution, but Flynn keeps it intact by manipulating the system again. Sark reaches the MCP's core on a shuttle carrying captured programs deemed powerful or useful. While the MCP attempts to absorb these programs, Tron, who is still alive, confronts Sark and critically injures him, prompting the MCP to give Sark all its functions. Realizing that his ability to manipulate the system might give Tron an opening, Flynn leaps into the beam of the MCP, distracting it. Seeing a break in the MCP's shield, Tron attacks through the gap and destroys the MCP and erases Sark, ending the MCP's control over the system, releasing all lockouts on computer access, and allowing the captured programs to communicate with users again. Flynn reappears in the real world, rematerialized at the terminal. A printed document proves Dillinger's guilt and Flynn as the original author of 'Space Paranoids'. The next morning, Dillinger enters his office to find the MCP deactivated and his theft exposed. Flynn is subsequently promoted to CEO of ENCOM and is happily greeted by Alan and Lora as their new boss.

Brothers Randolph and Mortimer Duke own a commodities brokerage firm, Duke & Duke Commodity Brokers, in Philadelphia, Pennsylvania. They witness an encounter between their managing director—the well-mannered and educated Louis Winthorpe III, engaged to the Dukes' grandniece Penelope Witherspoon—and poor black street hustler Billy Ray Valentine. Valentine is arrested at Winthorpe's insistence after the latter assumes he is being robbed. Holding opposing views on the issue of nature versus nurture, the Dukes make a wager and agree to conduct an experiment to observe the results of switching the lives of Valentine and Winthorpe, two people in contrasting social strata. Winthorpe is framed as a thief, drug dealer, and philanderer by Clarence Beeks, a man on the Dukes' payroll. He is fired from Duke & Duke, his bank accounts are frozen, he is denied entry to his Duke-owned home, and is ostracized by Penelope and his friends. Winthorpe is befriended by Ophelia, a prostitute who helps him in exchange for a financial reward once he is exonerated to secure her own retirement. The Dukes post bail for Valentine, install him in Winthorpe's former job, and grant him use of Winthorpe's home, including the service of his butler Coleman. Valentine becomes well versed in the business, using his street smarts to achieve success, and begins to act in a well-mannered way. During the firm's Christmas party, Winthorpe plants drugs in Valentine's desk, attempting to frame him, and brandishes a gun to escape. Later, the Dukes discuss their experiment and settle their wager for $1. They plot to return Valentine to the streets, but have no intention of taking back Winthorpe. Valentine overhears the conversation and seeks out Winthorpe, who has attempted suicide by overdosing. Valentine, Ophelia and Coleman nurse him back to health and inform him of the experiment. Watching a television news broadcast, they learn that Beeks is transporting a secret United States Department of Agriculture report on orange crop forecasts. Remembering large payments made to Beeks by the Dukes, Winthorpe and Valentine decide to foil the brothers' plan to obtain the report early and use it to corner the market on frozen concentrated orange juice. On New Year's Eve, the four board Beeks's train in disguise, intending to switch the original report with a forgery that predicts low orange crop yields. Beeks uncovers their scheme and attempts to kill them, but is knocked unconscious by a gorilla being transported on the train. The four dress him in a gorilla suit and cage him with the real gorilla. They deliver the forged report to the Dukes in Beeks's place and collect the payment intended for him. After sharing a kiss with Ophelia, Winthorpe travels to New York City with Valentine, pooling the money with the life savings of Ophelia and Coleman to carry out their plan. On the commodities trading floor, the Dukes invest heavily in buying frozen concentrated orange juice futures contracts, legally committing themselves to purchase the commodity at a later date. Other traders follow their lead, driving the price up; Valentine and Winthorpe naked-short-sell juice futures contracts at the inflated price. Following the broadcast of the actual crop report and its prediction of a normal harvest, the price of juice futures plummets. As the traders frantically sell their futures, Valentine and Winthorpe buy at the lower price from everyone except the Dukes, fulfilling the contracts they had short-sold earlier and turning an immense profit. After the closing bell, Valentine and Winthorpe explain to the Dukes that they made a wager on whether they could get rich and make the Dukes poor at the same time, and Winthorpe pays Valentine his winnings of $1. When the Dukes prove unable to pay the $394 million required to satisfy their margin call, the exchange manager orders their seats sold and their corporate and personal assets confiscated, effectively bankrupting them. Randolph collapses holding his chest and Mortimer shouts at the others, demanding the floor be reopened in a futile plea to recoup their losses. The now-wealthy Valentine, Winthorpe, Ophelia, and Coleman vacation on a luxurious tropical beach, while Beeks and the gorilla are loaded onto a ship bound for Africa.

In the Spanish countryside, a middle-aged man named Héctor and his wife Clara live in a home that they are renovating. Héctor scans the forest behind their house with binoculars and sees a young woman take off her T-shirt, exposing her breasts. When his wife goes shopping, he investigates and finds the woman on the ground, naked and unconscious. He is stabbed in the arm by a mysterious man with bloody bandages on his face. Fleeing and breaking into a mysterious nearby building, Héctor contacts a scientist by walkie-talkie, who warns him of the bandaged man and guides him to his location, promising safety. With the bandaged man just outside, the scientist convinces Héctor to hide inside a large mechanical device. When Héctor leaves the machine, he discovers that he has traveled approximately an hour back in time. The scientist explains that the machine is an experimental time machine and refers to Héctor as "Héctor 2". The scientist tells Héctor 2 that they need to stay where they are and let events unfold. Despite the scientist's warning, Héctor 2 drives off in a car, passing a cyclist, who he recognizes as the woman Héctor 1 saw in the forest. Héctor 2 chases the woman, only to be run off the road by a van, cutting his head, which he wraps using the bandage from his arm wound. He then realizes the bandaged man from before is himself. The woman approaches to see if he is all right. Héctor 2 replicates events by making the woman undress in view of Héctor 1. When she runs away, Héctor 2 catches her, inadvertently knocking her out. Héctor 2 lays her out naked on the ground and then stabs Hector 1 in the arm when he arrives. The woman escapes. Héctor 2 returns to his home, where he hears a scream and chases a woman through his house and onto the roof. When Héctor 2 attempts to grab her, she slips and falls to her death. Seeing the body from the roof, Héctor 2 is horrified, believing he has killed his own wife. Héctor 2 contacts the scientist over the walkie-talkie and convinces him to lure Héctor 1 to the lab with warnings that he is being pursued. Driving to the lab, Héctor 2 insists that he must travel back one more time, despite the scientist revealing that there is a Héctor 3, who told him he must stop Héctor 2 from doing just that. After removing his bandages, Héctor 2 convinces the scientist to send him back several seconds before Héctor 2 initially appears. This causes him to become Héctor 3, who uses a van to run Héctor 2 off the road, but crashes as well, knocking himself out. Upon waking, Héctor 3 informs the scientist he has failed to stop Héctor 2. Héctor 3 encounters the woman again, startling her into screaming, though she does not recognize him as her assailant. Since Héctor 2 has heard her scream, Héctor 3 and the woman flee to Héctor's house. They become separated. Héctor 3 finds and hides his wife, then realizes what has to happen / will happen / has already happened. He finds the woman, cuts her ponytail off, gives her his wife's coat and tells her to hide upstairs. Héctor 2 chases her onto the roof. Héctor 3 sits on his lawn with his wife as Héctor 2 accidentally kills the woman, then drives off – heading back to the lab to become Héctor 3. Emergency vehicles are heard approaching in the distance.

In February 1958, the Second Cross-Winter Expedition for the Japanese Antarctic Surveying Team rides on the icebreaker Sōya to take over from the 11-man First Cross-Winter Expedition. The First Cross-Winter Expedition retreats by helicopter, leaving 15 Sakhalin huskies chained up at the Showa Base for the next Expedition. Due to the extreme weather conditions, Sōya can not get near enough to the base and it is decided not to proceed with the handover, leaving the base unmanned. The team is worried about the dogs, as the weather is extremely cold and only one week of food for the dogs has been left. They wish to rescue them but in the end are unable to, due to a shortage of fuel and drinking water. Eight of the fifteen sled dogs manage to break loose from their chains (Riki, Anko, Shiro, Jakku, Deri, Kuma, Taro, and Jiro), while the other seven starve. As the eight journey across the frozen wilderness, they are forced to survive by hunting penguins and seals on the ice shelves and even by eating seal excrement. As the months pass, most die or disappear. Riki is fatally injured by a killer whale while trying to protect Taro and Jiro. Anko and Deri fall through the ice and drown in the freezing waters. Shiro falls off a cliff to his death, and Jakku and Kuma disappear in the wilderness. Eleven months later, on 14 January 1959, Kitagawa, one of the dog handlers in the first expedition, returns with the Third Cross-Winter Expedition, wanting to bury his beloved dogs. He, along with the two dog-handlers Ushioda and Ochi, recover the frozen corpses of the seven chained dogs, but are surprised to discover that eight others have broken loose. To everyone's surprise, they are greeted warmly at the base by Taro and Jiro, brothers who were born in Antarctica. It is still unknown how and why they survived, because an average husky can only live in such conditions for about one month. In the movie, the director used the data available, together with his imagination, to reconstruct how the dogs struggled with the elements and survived.