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Bob Munro, an executive at the large soda company Pure Vibe, struggles with the whims of his self-absorbed boss Todd Mallory. His family — materialistic wife Jamie and their teenage children, sharp-tongued Cassie and self-confident Carl — are also demanding, and he had promised them a vacation in Hawaii. During an event Todd hosted, Cassie's friend Gretchen insults Todd and stains his suit with a soda. Todd, who sought to acquire the Alpine Soda company in Boulder, Colorado, blames Bob for the incident and threatens to fire him if he does not promote the takeover. Todd's demand forces the Munros to cancel their vacation. Bob, concealing the real reason for not going to Hawaii, rents an RV and tells his family that they are traveling to the Rockies; he plans to make a detour in Colorado to secretly attend the meeting in Boulder.
The trip is marked by numerous mishaps. Bob's inexperience in handling the large vehicle results in him colliding with various obstacles and damaging the parking brake. During a stopover in Nevada, he must fix an unsavory clog in the toilet tank. The family also fumigates the RV with stink bombs to drive out intrusive raccoons. During the trip, the Munros have several encounters with the Gornickes, a good-natured but exhausting family who live in their own RV, consisting of dad Travis, mobile sales rep and cosmetologist mom Mary Jo, and their three children, Earl, Moon, and Billy.
As they approach Colorado, Carl tries playing basketball with some older boys, but Bob makes it awkward and embarrasses him. The two go for a walk in the woods and somehow bond over the future of Carl's heavyweight career. Later, the Munros reconnect as a family and enjoy the beauty of their surroundings. Nearing Boulder, Bob fakes an upset stomach and sends the rest of the family on a hike before meeting with the Alpine Soda owners. The meeting is promising, but on the way back, Bob gets stuck in a traffic jam, forcing him to take the RV through a treacherous four-wheel-drive trail; he returns to his family with a badly battered vehicle.
Todd calls Bob and demands that he return the next day to repeat the presentation to the entire Alpine Soda staff. However, the RV's parking brake fails again and sends it rolling into a mountain lake. Bob is forced to confess the true purpose of the trip to his family, explaining that he fears losing his job and their standard of living. Taken aback, the family refuses to accompany him, leaving him to make his way to the meeting alone on a bike recovered from the RV. The Gornickes appear and pick up the Munros. Along the way, the two families bond; Bob spots them together from a distance on the road. Deciding his family is more important than his job, he catches up to them and reconciles with everyone. In return, the Gornickes drive him the rest of the way to the meeting.
At the Alpine Soda headquarters, Bob begins to pitch the takeover to the staff; halfway through, he has an epiphany and encourages their independence, aware that Todd would ruin the brand. In retaliation, Todd fires Bob; Carl angrily throws to the ground and tackles him, making him a hero. Then Bob retorts that he quits anyway. On their way home in the sodden and battered RV, the Munros are stopped by a police officer on behalf of the Alpine Soda owners, who offer Bob a job overseeing their company's expansion. The RV's parking brake fails again, flattening the police car and the Alpine Soda company owners' car.
During the credits, the two families are shown dancing and singing along to " Route 66 ".